(i) Chromosomes are moved to spindle equator during the metaphase stage. (ii) Centromere splits and chromatids separate during the anaphase. (iii) Pairing between homologous chromosomes takes place during zygotene stage of prophase I in meiosis. (iv) Crossing over between homologous chromosomes takes place during the pachytene stage of prophase I in meiosis.
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